1. "puffed with Darwinian pieties for killing" (p. 659)
2. My guess is that it is in some way related to Darwinism. I think this means that because of survival of the fittest this creature is suppose to do this or naturally does this in an effort to survive.
3. Definition: Having or showing Duty to God or any religious fulfillment. Dutifulness
4. What this phrase is saying with this word is not in reference to god but instead to a dutiful obligation to what they are doing.
Dictionary.com
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