1) Thence to a lightness and by this declension [...]"(1333,147)
2) My Guess: The word looks similar to decline or regression, it makes sense when speaking of Hamlet's current decline in character.
3) Definition: "The inflection of nouns, pronouns, and adjectives for categories such as case and number" (dictionary.com)
4) I guess Polonius is commenting on how Hamlet now speaks in his madness, how it differs from his normal speech.
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